In ecology, the theory of alternative stable states predicts that ecosystems can exist under multiple "states". These alternative states are non-transitory and therefore considered stable over ecologically-relevant timescales. Ecosystems may transition from one stable state to another, in what is known as a state shift, when perturbed. Due to ecological feedbacks, ecosystems display resistance to state shifts and therefore tend to remain in one state unless perturbations are large enough. Multiple states may persist under equal environmental conditions, a phenomenon known as hysteresis. Alternative stable state theory suggests that discrete states are separated by ecological thresholds, in contrast to ecosystems which change smoothly and continuously along an environmental gradient.